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What's the difference between the play button and the button to the right of it? They seem to do the same thing

(+1)

The button to the right of it is to queue an action, quite helpful actually

(+2)

To clarify - the play button (Start Action) puts it at the front of the queue, so you start on it immediately. The second button (Queue action) adds it to the back of the queue, so you'll start it when you finish all the other actions in your queue.